Notes
Enzyme Naming Conventions
Sections
Overview
ENZYME NOMENCLATURE
- Derived from substrate or chemical reaction the enzyme catalyzes
- Most enzyme names have suffix "–ase"
Isozymes (isoenzymes)
- Enzymes that catalyze same reaction but have different amino acid sequences (different proteins)
ENZYMES REQUIRING COFACTORS
Apoenzyme
- Enzyme without the cofactor.
Holoenzyme
- Enzyme and cofactor together
6 classes of enzymes
Oxidoreductases
- Catalyze electron transfer reactions (oxidation/reduction reactions)
- A- + B → A + B-
Transferases
- Catalyze transfer of functional groups to a molecule
- A-B + PO4 → A-B-PO4
Hydrolases
- Catalyze breakage of bonds by hydrolysis (addition of water)
- A-B-C + H2O → A-B + C
Lyases
- Catalyze bond cleavage reactions that are not oxidation or hydrolysis reactions
- A-B-C → A-B + C
Isomerases
- Catalyze rearrangement reactions
- A-B-C → A-C-B
Ligases
- Catalyze reactions in which covalent bonds join two molecules
- A-D + B-C→ A-D-B-C
Full-Length Text
- Here we will learn about the naming conventions of enzymes: their characteristic activity typically determines their nomenclature and classification.
- Start a table.
- Denote that enzyme names are derived from the substrate or the chemical reaction the enzyme catalyzes.
- Denote that most enzyme names have the suffix "–ase."
- Denote that isozymes (aka isoenzymes) are enzymes that catalyze the same reaction but have different amino acids sequences (so they are different proteins).
- Denote that for enzymes requiring cofactors:
- An apoenzyme is an enzyme without the cofactor.
- A holoenzyme is the enzyme and cofactor together.
Let's spend the remainder of our time learning the six classes of enzymes that cover every type of enzyme-catalyzed reaction.
- Denote that they are:
- Oxidoreductases
- Transferases
- Hydrolases
- Lyases
- Isomerases
- Ligases
Let's start with oxidoreductases.
- Write that oxidoreductases catalyze electron transfer reactions (oxidation or reduction reactions).
- As an example, create an equation wherein oxidoreductase catalyzes the transfer of electrons from molecule X to molecule Y.
- List dehydrogenases as a specific type of oxidoreductase.
- As a clinical correlation, write that alcohol dehydrogenase metabolizes alcohol via oxidation, thus alcohol affects individuals differently based on their level and isoform of alcohol dehydrogenase.
- Now, write that transferases catalyze the transfer of functional groups to a molecule.
- To see how, draw an amorphous protein.
- Then, show draw that protein with a group added to it.
- Draw reversible arrows with a group above them to represent how transferases add functional groups to a molecule, such as a protein or a lipid.
- List phosphotransferases as a type of transferase.
- As a clinical correlation, write that the transferase phosphofructokinase transfers phosphate groups to substrates.
- Phosphofructokinase deficiency results in a glycogen storage disease (Tauri's disease) wherein exercise causes patients to develop muscle cramps, weakness, and dark urine (from hemolytic anemia).
- Now, write that hydrolases catalyze the breakage of bonds by hydrolysis (the addition of water).
- To illustrate this, draw a drop of water breaking the bond between two molecules.
- Hydrolysis is one of the most common physiological ways of breaking bonds.
- List nucleases (which cut nucleic acids) and proteases (which cut proteins) as types of hydrolases.
- As an experimental correlation, write that endonucleases (called restriction enzymes) cut specific sequences of DNA and are used for DNA modification and molecular cloning.
- Now, write that lyases catalayze bond cleavage reactions that are not oxidation or hydrolysis reactions.
- Draw a pair of scissors cutting the bond between two molecules to illustrate the action of lyases.
- List carboxylases and decarboxylases as examples of lyases.
- As an ecological correlation, RuBisCO (ribulose-1,5-bisphosphate carboxylase/oxygenase) is a plant lyase that cleaves a 5-carbon sugar in the first steps of carbon fixation; it is one of the most abundant and important proteins on earth.
- Next, write that isomerases catalyze rearrangement reactions.
- As an abstraction, use several shapes to create a cat
- Then, show that isomerases rearrange those shapes to form a bird, much like they rearrange atoms to turn one molecule into another.
- As an example, list triose phosphate isomerase, which converts dihydroxyacteone phosphate to D-glyceraldehyde 3-phosphate during glycolysis.
- Now, write that ligases catalyze reactions in which covalent bonds join two molecules.
- As another abstraction, draw a large snowball, then a small one.
- Next, stack to them to show that ligases join two molecules together.
- As an example, list DNA ligase, which joins two strands of DNA together to catalyze the formation of a phosphodiester bond.